ANNEX II
MARINE ECOSYSTEMS – HABITAT TYPES AND GROUPS OF HABITAT TYPES REFERRED TO IN ARTICLE 5(1) AND (2)
The list below includes the marine habitat types referred to in Article 5(1) and (2), as well as seven groups of those habitat types, namely 1) seagrass beds, 2) macroalgal forests, 3) shellfish beds, 4) maerl beds, 5) sponge, coral and coralligenous beds, 6) vents and seeps and 7) soft sediments (not deeper than 1 000 metres of depth). The relation with the habitat types listed in Annex I to Directive 92/43/EEC is also presented.
The classification of marine habitat types used, differentiated by marine biogeographical regions, is made according to the European nature information system (EUNIS), as revised for the marine habitats typology in 2022 by the EEA. The information on the related habitats listed in Annex I to Directive 92/43/EEC is based on the crosswalk published by the EEA in 2021 ( 1 ) .
1. Group 1: Seagrass beds
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MA522
Seagrass beds on Atlantic littoral sand
1140; 1160
MA623
Seagrass beds on Atlantic littoral mud
1140; 1160
MB522
Seagrass beds on Atlantic infralittoral sand
1110; 1150; 1160
Baltic Sea
MA332
Baltic hydrolittoral coarse sediment characterised by submerged vegetation
1130; 1160; 1610; 1620
MA432
Baltic hydrolittoral mixed sediment characterised by submerged vegetation
1130; 1140; 1160; 1610
MA532
Baltic hydrolittoral sand characterised by submerged rooted plants
1130; 1140; 1160; 1610
MA632
Baltic hydrolittoral mud dominated by submerged rooted plants
1130; 1140; 1160; 1650
MB332
Baltic infralittoral coarse sediment characterised by submerged rooted plants
1110; 1160
MB432
Baltic infralittoral mixed sediment characterised by submerged rooted plants
1110; 1160; 1650
MB532
Baltic infralittoral sand characterised by submerged rooted plants
1110; 1130; 1150; 1160
MB632
Baltic infralittoral mud sediment characterised by submerged rooted plants
1130; 1150; 1160; 1650
Black Sea
MB546
Seagrass and rhizomatous algal meadows in Black Sea freshwater influenced infralittoral muddy sands
1110; 1130; 1160
MB547
Black Sea seagrass meadows on moderately exposed upper infralittoral clean sands
1110; 1160
MB548
Black Sea seagrass meadows on lower infralittoral sands
1110; 1160
Mediterranean Sea
MB252
Biocenosis of Posidonia oceanica
1120
MB2521
Ecomorphosis of striped Posidonia oceanica meadows
1120; 1130; 1160
MB2522
Ecomorphosis of ‘barrier-reef’ Posidonia oceanica meadows
1120; 1130; 1160
MB2523
Facies of dead ‘mattes’ of Posidonia oceanica without much epiflora
1120; 1130; 1160
MB2524
Association with Caulerpa prolifera on Posidonia beds
1120; 1130; 1160
MB5521
Association with Cymodocea nodosa on well sorted fine sands
1110; 1130; 1160
MB5534
Association with Cymodocea nodosa on superficial muddy sands in sheltered waters
1110; 1130; 1160
MB5535
Association with Zostera noltei on superficial muddy sands in sheltered waters
1110; 1130; 1160
MB5541
Association with Ruppia cirrhosa and/or Ruppia maritima on sand
1110; 1130; 1160
MB5544
Association with Zostera noltei in euryhaline and eurythermal environment on sand
1110; 1130; 1160
MB5545
Association with Zostera marina in euryhaline and eurythermal environment
1110; 1130; 1160
2. Group 2: Macroalgal forests
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MA123
Seaweed communities on full salinity Atlantic littoral rock
1160; 1170; 1130
MA125
Fucoids on variable salinity Atlantic littoral rock
1170; 1130
MB121
Kelp and seaweed communities on Atlantic infralittoral rock
1170; 1160
MB123
Kelp and seaweed communities on sediment-affected or disturbed Atlantic infralittoral rock
1170; 1160
MB124
Kelp communities on variable salinity Atlantic infralittoral rock
1170; 1130; 1160
MB321
Kelp and seaweed communities on Atlantic infralittoral coarse sediment
1160
MB521
Kelp and seaweed communities on Atlantic infralittoral sand
1160
MB621
Vegetated communities on Atlantic infralittoral mud
1160
Baltic Sea
MA131
Baltic hydrolittoral rock and boulders characterised by perennial algae
1160; 1170; 1130; 1610; 1620
MB131
Perennial algae on Baltic infralittoral rock and boulders
1170; 1160
MB232
Baltic infralittoral bottoms characterised by shell gravel
1160; 1110
MB333
Baltic infralittoral coarse sediment characterised by perennial algae
1110; 1160
MB433
Baltic infralittoral mixed sediment characterised by perennial algae
1110; 1130; 1160; 1170
Black Sea
MB144
Mytilid-dominated Black Sea exposed upper infralittoral rock with fucales
1170; 1160
MB149
Mytilid-dominated Black Sea moderately exposed upper infralittoral rock with fucales
1170; 1160
MB14A
Fucales and other algae on Black Sea sheltered upper infralittoral rock, well illuminated
1170; 1160
Mediterranean Sea
MA1548
Association with Fucus virsoides
1160; 1170
MB1512
Association with Cystoseira tamariscifolia and Saccorhiza polyschides
1170; 1160
MB1513
Association with Cystoseira amentacea (var. amentacea , var. stricta , var. spicata )
1170; 1160
MB151F
Association with Cystoseira brachycarpa
1170; 1160
MB151G
Association with Cystoseira crinita
1170; 1160
MB151H
Association with Cystoseira crinitophylla
1170; 1160
MB151J
Association with Cystoseira sauvageauana
1170; 1160
MB151K
Association with Cystoseira spinosa
1170; 1160
MB151L
Association with Sargassum vulgare
1170; 1160
MB151M
Association with Dictyopteris polypodioides
1170; 1160
MB151W
Association with Cystoseira compressa
1170; 1160
MB1524
Association with Cystoseira barbata
1170; 1160
MC1511
Association with Cystoseira zosteroides
1170; 1160
MC1512
Association with Cystoseira usneoides
1170; 1160
MC1513
Association with Cystoseira dubia
1170; 1160
MC1514
Association with Cystoseira corniculata
1170; 1160
MC1515
Association with Sargassum spp.
1170; 1160
MC1518
Association with Laminaria ochroleuca
1170; 1160
MC3517
Association with Laminaria rodriguezii on detritic beds
1160
3. Group 3: Shellfish beds
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MA122
Mytilus edulis and/or barnacle communities on wave-exposed Atlantic littoral rock
1160; 1170
MA124
Mussel and/or barnacle communities with seaweeds on Atlantic littoral rock
1160; 1170
MA227
Bivalve reefs in the Atlantic littoral zone
1170; 1140
MB222
Bivalve reefs in the Atlantic infralittoral zone
1170; 1130; 1160
MC223
Bivalve reefs in the Atlantic circalittoral zone
1170
Baltic Sea
MB231
Baltic infralittoral bottoms dominated by epibenthic bivalves
1170; 1160
MC231
Baltic circalittoral bottoms dominated by epibenthic bivalves
1170; 1160; 1110
MD231
Baltic offshore circalittoral biogenic bottoms characterised by epibenthic bivalves
1170
MD232
Baltic offshore circalittoral shell gravel bottoms characterised by bivalves
1170
MD431
Baltic offshore circalittoral mixed bottoms characterised by macroscopic epibenthic biotic structures
MD531
Baltic offshore circalittoral sand characterised by macroscopic epibenthic biotic structures
MD631
Baltic offshore circalittoral mud characterised by epibenthic bivalves
Black Sea
MB141
Invertebrate-dominated Black Sea lower infralittoral rock
1170
MB143
Mytilid-dominated Black Sea exposed upper infralittoral rock with foliose algae (no Fucales)
1170; 1160
MB148
Mytilid-dominated Black Sea moderately exposed upper infralittoral rock with foliose algae (other than Fucales)
1170; 1160
MB242
Mussel beds in the Black Sea infralittoral zone
1170; 1130; 1160
MB243
Oyster reefs on Black Sea lower infralittoral rock
1170
MB642
Black Sea infralittoral terrigenous muds
1160
MC141
Invertebrate-dominated Black Sea circalittoral rock
1170
MC241
Mussel beds on Black Sea circalittoral terrigenous muds
1170
MC645
Black Sea lower circalittoral mud
Mediterranean Sea
MA1544
Facies with Mytilus galloprovincialis in waters enriched in organic matter
1160; 1170
MB1514
Facies with Mytilus galloprovincialis
1170; 1160
Mediterranean infralittoral oyster beds
Mediterranean circalittoral oyster beds
4. Group 4: Maerl beds
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MB322
Maerl beds on Atlantic infralittoral coarse sediment
1110; 1160
MB421
Maerl beds on Atlantic infralittoral mixed sediment
1110; 1160
MB622
Maerl beds on Atlantic infralittoral muddy sediment
1110; 1160
Mediterranean Sea
MB3511
Association with rhodolithes in coarse sands and fine gravels mixed by waves
1110; 1160
MB3521
Association with rhodolithes in coarse sands and fine gravels under the influence of bottom currents
1110; 1160
MB3522
Association with maerl (= Association with Lithothamnion corallioides and Phymatolithon calcareum ) on Mediterranean coarse sands and gravel
1110; 1160
MC3521
Association with rhodolithes on coastal detritic bottoms
1110
MC3523
Association with maerl ( Lithothamnion corallioides and Phymatholithon calcareum ) on coastal dendritic bottoms
1110
5. Group 5: Sponge, coral and coralligenous beds
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MC121
Faunal turf communities on Atlantic circalittoral rock
1170
MC124
Faunal communities on variable salinity Atlantic circalittoral rock
1170; 1130
MC126
Communities of Atlantic circalittoral caves and overhangs
8330; 1170
MC222
Cold water coral reefs in the Atlantic circalittoral zone
1170
MD121
Sponge communities on Atlantic offshore circalittoral rock
1170
MD221
Cold water coral reefs in the Atlantic offshore circalittoral zone
1170
ME122
Sponge communities on Atlantic upper bathyal rock
1170
ME123
Mixed cold water coral communities on Atlantic upper bathyal rock
1170
ME221
Atlantic upper bathyal cold water coral reef
1170
ME322
Mixed cold water coral community on Atlantic upper bathyal coarse sediment
ME324
Sponge aggregation on Atlantic upper bathyal coarse sediment
ME422
Sponge aggregation on Atlantic upper bathyal mixed sediment
ME623
Sponge aggregation on Atlantic upper bathyal mud
ME624
Erect coral field on Atlantic upper bathyal mud
MF121
Mixed cold water coral community on Atlantic lower bathyal rock
1170
MF221
Atlantic lower bathyal cold water coral reef
1170
MF321
Mixed cold water coral community on Atlantic lower bathyal coarse sediment
MF622
Sponge aggregation on Atlantic lower bathyal mud
MF623
Erect coral field on Atlantic lower bathyal mud
Baltic Sea
MB138
Baltic infralittoral rock and boulders characterized by epibenthic sponges
1170; 1160
MB43A
Baltic infralittoral mixed sediment characterized by epibenthic sponges (Porifera)
1160; 1170
MC133
Baltic circalittoral rock and boulders characterized by epibenthic cnidarians
1170; 1160
MC136
Baltic circalittoral rock and boulders characterized by epibenthic sponges
1170; 1160
MC433
Baltic circalittoral mixed sediment characterized by epibenthic cnidarians
1160; 1170
MC436
Baltic circalittoral mixed sediment characterized by epibenthic sponges
1160
Black Sea
MD24
Black Sea offshore circalittoral biogenic habitats
1170
ME14
Black Sea upper bathyal rock
1170
ME24
Black Sea upper bathyal biogenic habitat
1170
MF14
Black Sea lower bathyal rock
1170
Mediterranean Sea
MB151E
Facies with Cladocora caespitosa
1170; 1160
MB151Q
Facies with Astroides calycularis
1170; 1160
MB151α
Facies and association of coralligenous biocenosis (in enclave)
1170; 1160
MC1519
Facies with Eunicella cavolini
1170; 1160
MC151A
Facies with Eunicella singularis
1170; 1160
MC151B
Facies with Paramuricea clavata
1170; 1160
MC151E
Facies with Leptogorgia sarmentosa
1170; 1160
MC151F
Facies with Anthipatella subpinnata and sparse red algae
1170; 1160
MC151G
Facies with massive sponges and sparse red algae
1170; 1160
MC1522
Facies with Corallium rubrum
8330; 1170
MC1523
Facies with Leptopsammia pruvoti
8330; 1170
MC251
Coralligenous platforms
1170
MC6514
Facies of sticky muds with Alcyonium palmatum and Parastichopus regalis on circalittoral mud
1160
MD151
Biocenosis of Mediterranean shelf-edge rock
1170
MD25
Mediterranean offshore circalittoral biogenic habitats
1170
MD6512
Facies of sticky muds with Alcyonium palmatum and Parastichopus regalis on lower circalittoral mud
ME1511
Mediterranean upper bathyal Lophelia pertusa reefs
1170
ME1512
Mediterranean upper bathyal Madrepora oculata reefs
1170
ME1513
Mediterranean upper bathyal Madrepora oculata and Lophelia pertusa reefs
1170
ME6514
Mediterranean upper bathyal facies of with Pheronema carpenteri
MF1511
Mediterranean lower bathyal Lophelia pertusa reefs
1170
MF1512
Mediterranean lower bathyal Madrepora oculata reefs
1170
MF1513
Mediterranean lower bathyal Madrepora oculata and Lophelia pertusa reefs
1170
MF6511
Mediterranean lower bathyal facies of sandy muds with Thenea muricata
MF6513
Mediterranean lower bathyal facies of compact muds with Isidella elongata
6. Group 6: Vents and seeps
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MB128
Vents and seeps in Atlantic infralittoral rock
1170; 1160; 1180
MB627
Vents and seeps in Atlantic infralittoral mud
1130; 1160
MC127
Vents and seeps in Atlantic circalittoral rock
1170; 1180
MC622
Vents and seeps in Atlantic circalittoral mud
1160
MD122
Vents and seeps on Atlantic offshore circalittoral rock
1170
MD622
Vents and seeps in Atlantic offshore circalittoral mud
7. Group 7: Soft sediments (not deeper than 1 000 metres of depth)
EUNIS code
EUNIS habitat type name
Related habitat type code as referred to in Annex I to Directive 92/43/EEC
Atlantic
MA32
Atlantic littoral coarse sediment
1130; 1160
MA42
Atlantic littoral mixed sediment
1130; 1140; 1160
MA52
Atlantic littoral sand
1130; 1140; 1160
MA62
Atlantic littoral mud
1130; 1140; 1160
MB32
Atlantic infralittoral coarse sediment
1110; 1130; 1160
MB42
Atlantic infralittoral mixed sediment
1110; 1130; 1150; 1160
MB52
Atlantic infralittoral sand
1110; 1130; 1150; 1160
MB62
Atlantic infralittoral mud
1110; 1130; 1160
MC32
Atlantic circalittoral coarse sediment
1110; 1160
MC42
Atlantic circalittoral mixed sediment
1110; 1160
MC52
Atlantic circalittoral sand
1110; 1160
MC62
Atlantic circalittoral mud
1160
MD32
Atlantic offshore circalittoral coarse sediment
MD42
Atlantic offshore circalittoral mixed sediment
MD52
Atlantic offshore circalittoral sand
MD62
Atlantic offshore circalittoral mud
ME32
Atlantic upper bathyal coarse sediment
ME42
Atlantic upper bathyal mixed sediment
ME52
Atlantic upper bathyal sand
ME62
Atlantic upper bathyal mud
MF32
Atlantic lower bathyal coarse sediment
MF42
Atlantic lower bathyal mixed sediment
MF52
Atlantic lower bathyal sand
MF62
Atlantic lower bathyal mud
Baltic Sea
MA33
Baltic hydrolittoral coarse sediment
1130; 1160; 1610; 1620
MA43
Baltic hydrolittoral mixed sediment
1130; 1140; 1160; 1610
MA53
Baltic hydrolittoral sand
1130; 1140; 1160; 1610
MA63
Baltic hydrolittoral mud
1130; 1140; 1160; 1650
MB33
Baltic infralittoral coarse sediment
1110; 1150; 1160
MB43
Baltic infralittoral mixed sediment
1110; 1130; 1150; 1160; 1170; 1650
MB53
Baltic infralittoral sand
1110; 1130; 1150; 1160
MB63
Baltic infralittoral mud
1130; 1150; 1160; 1650
MC33
Baltic circalittoral coarse sediment
1110; 1160
MC43
Baltic circalittoral mixed sediment
1160; 1170
MC53
Baltic circalittoral sand
1110; 1160
MC63
Baltic circalittoral mud
1160; 1650
MD33
Baltic offshore circalittoral coarse sediment
MD43
Baltic offshore circalittoral mixed sediment
MD53
Baltic offshore circalittoral sand
MD63
Baltic offshore circalittoral mud
Black Sea
MA34
Black Sea littoral coarse sediment
1160
MA44
Black Sea littoral mixed sediment
1130; 1140; 1160
MA54
Black Sea littoral sand
1130; 1140; 1160
MA64
Black Sea littoral mud
1130; 1140; 1160
MB34
Black Sea infralittoral coarse sediment
1110; 1160
MB44
Black Sea infralittoral mixed sediment
1110; 1170
MB54
Black Sea infralittoral sand
1110; 1130; 1160
MB64
Black Sea infralittoral mud
1130; 1160
MC34
Black Sea circalittoral coarse sediment
1160
MC44
Black Sea circalittoral mixed sediment
MC54
Black Sea circalittoral sand
1160
MC64
Black Sea circalittoral mud
1130; 1160
MD34
Black Sea offshore circalittoral coarse sediment
MD44
Black Sea offshore circalittoral mixed sediment
MD54
Black Sea offshore circalittoral sand
MD64
Black Sea offshore circalittoral mud
Mediterranean Sea
MA35
Mediterranean littoral coarse sediment
1160; 1130
MA45
Mediterranean littoral mixed sediment
1140; 1160
MA55
Mediterranean littoral sand
1130; 1140; 1160
MA65
Mediterranean littoral mud
1130; 1140; 1150; 1160
MB35
Mediterranean infralittoral coarse sediment
1110; 1160
MB45
Mediterranean infralittoral mixed sediment
MB55
Mediterranean infralittoral sand
1110; 1130; 1150; 1160
MB65
Mediterranean infralittoral mud
1130; 1150
MC35
Mediterranean circalittoral coarse sediment
1110; 1160
MC45
Mediterranean circalittoral mixed sediment
MC55
Mediterranean circalittoral sand
1110; 1160
MC65
Mediterranean circalittoral mud
1130; 1160
MD35
Mediterranean offshore circalittoral coarse sediment
MD45
Mediterranean offshore circalittoral mixed sediment
MD55
Mediterranean offshore circalittoral sand
MD65
Mediterranean offshore circalittoral mud
ME35
Mediterranean upper bathyal coarse sediment
ME45
Mediterranean upper bathyal mixed sediment
ME55
Mediterranean upper bathyal sand
ME65
Mediterranean upper bathyal mud
MF35
Mediterranean lower bathyal coarse sediment
MF45
Mediterranean lower bathyal mixed sediment
MF55
Mediterranean lower bathyal sand
MF65
Mediterranean lower bathyal mud
( 1 ) EUNIS marine habitat classification 2022. European Environment Agency https://www.eea.europa.eu/data-and-maps/data/eunis-habitat-classification-1 .